My general area of research interest is the genetic basis of adaptive trait evolution in natural populations, especially as it relates to the origin of new species via plant-pollinator coevolution. I use a combination of laboratory, greenhouse/growth chamber, 3D-printed flowers, and field experiments to study adaptation. My academic background is largely in molecular biology and biochemistry, but I have a lifelong interest in organismal biology. I collaborate with ecologists, physiologists, and evolutionary biologists in many of my research projects. I am no longer accepting new graduate students or postdocs.

Employment
Chair, Department of Biology, UW, 2011-
Washington Research Foundation Professor of Basic Biology, UW, 2005-7
Professor, Department of Biology, University of Washington, 2003-
Research Associate Professor, College of Forest Resources, University of Washington, 1995-2003
Research Associate Professor (adjunct), Botany Department, Univ. of Washington 1995-
Research Assistant Professor, College of Forest Resources, Univ. of Washington, 1994-5
Research Assistant Professor, Dept. of Biochemistry, Univ. of Washington, 1989-94
Helen Hay Whitney Postdoctoral Fellow (Milton P. Gordon, research director), 1984-87
National Science Foundation Graduate Fellow, 1980-83
American Heart Association/Louisiana Graduate Fellow, 1980-81
Graduate assistant, LSUMC (William C. Claycomb, research director), 1979-80