Ontogeny of Megalolaelaps colossus sp. nov.(Acari: Megalolaelapidae), an enigmatic symbiont of dung beetles (Coleoptera: Scarabaeidae) in Colombia, Cómbita-Heredia Orlando, Quintero-Gutiérrez Edwin Javier, and Klompen Hans, Systematic and Applied Acarology, Volume 23, p.1102–1124 (2018)